Conceptualization, J.L. and Z.C.; formal analysis, J.L. and J.D.; investigation, J.D., Q.T., C.Y. and Y.L.; methodology, J.L., J.Y. and Z.C.; project administration, J.L. and Z.C.; funding acquisition, J.L. and Z.C.; resources, J.L., J.Y. and Z.C.; supervision, J.L. and Z.C.; writing—original draft, J.L. and J.D.; writing—review and editing, J.L., J.Y. and Z.C. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (Nos. 21605009, 21275022), the Natural Science Foundation of Hunan Province (Nos. 2019JJ50651, 2025JJ50092) and the Scientific Research Fund of the Hunan Provincial Education Department (Nos. 21B0302, 24A0236).