Conceptualization, W.Q. and C.W.; methodology, B.W.; software, B.W.; investigation, B.W.; resources, J.H. and J.D.; data curation, W.Q., C.W. and Z.M.; writing—original draft preparation, B.W. and H.X.; writing—review and editing, W.Q., C.W. and Z.M.; supervision, J.H. and J.D.; project administration, W.Q., C.W.; funding acquisition, W.Q., C.W. and Z.M. All authors have read and agreed to the published version of the manuscript. This work was supported by the Natural Science Foundation of Hunan Province (Project Nos. 2020JJ5580; 2020JJ5563); the Scientific Research Foundation of Hunan Provincial Education Department (Contract No. 21B0328); the Scientific and Technological Plan Projects of Guangzhou City (No. 202103040001); the Fellowship of China Postdoctoral Science Foundation (No. 2020M672670).